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I discussed this matter while ago with one frien who is using screen reader and speech synthesizer. He did some testing with my Pligg site and found out that browsing the site and using user features was on a quite normal accessibility level. He said that it wasn't the best user experience but there wasn't any bigger problems with it.
I think that if your template is well structured there shouldn't be any big problems. Just remember to use Ajax and Javascript visely and it's all good.
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